Ph.D., Ecology and Evolutionary Biology, Iowa State University
Dissertation: "Patterns and processes of spatial genetic structure in a mobile and continuously distributed species, the bobcat (Lynx rufus)"
M.S., Zoology with specialization in Ecology, Evolution, and Conservation Biology, University of Hawaii
Thesis: "Current and historical genetic diversity and structure in the endangered Hawaii Akepa"
B.S., Environmental Science, University of Dubuque