This course is an introduction to the ecology, evolution, and natural history of the Amazon rainforest, Andean cloud forest, and Galapagos Islands. At the same time, the course exposes students to some of the customs and diverse cultures of the Ecuadorian people. We will review basic principles of evolution, ecology, and natural history as they pertain to both the rainforest, cloud forest (e.g. productivity, nutrient cycling, succession, and coevolution), and Galapagos Islands (e.g. natural selection, speciation, colonization, and island biogeography).