Ph.D., Biology, Utah State University, 1980
Dissertation: "Antipredator behavior and parental care in the American Avocet and Black-necked Stilt (Aves: Recurvirostridae)."
M.S., Zoology, Utah State University, 1976 (University Research Fellow, 1973–4)
Thesis: "Adaptive aspects of the vocal repertoire of the Willet."
B.A., magna cum laude, Biology, Luther College, 1973
Senior Thesis: "Bird life in a boreal sphagnum bog at the Audubon Camp of Wisconsin."